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 PUBLICATIONS

 -= FACULTY =-
James Boone
Angela Bryan
M. Emery Thompson
Keith Hunley
Steve Gangestad
Hillard Kaplan
Jane Lancaster
Geoffrey Miller
Martin Muller
Sherry Nelson
Osbjorn Pearson
Randy Thornhill
Paul Watson


Randy Thornhill
rthorn@unm.edu
Behavioral Ecology / Evolutionary Biology

personal website




     BOOKS     SELECTED PUBLICATIONS
    in press
  • In press S.W. Gangestad and R. Thornhill. Human estrus. Proceedings of the Royal Society of London B.

  • 2007
  • 2007. R. Thornhill and C.L. Fincher. What is the relevance of attachment and life history to political values? Evolution and Human Behavior 28:215-222.
  • 2007. R. Thornhill. The importance of developmental biology to evolutionary biology and vice versa. Pp. 203-209 in The Evolution of Mind: Fundamental Questions and Controversies, S.W. Gangestad and J.A. Simpson, eds. Guilford Publications, Inc., New York, NY.
  • 2007. R. Thornhill. The evolution of woman’s estrus, extended sexuality and concealed ovulation and their implications for human sexuality research. Pp. 391-396 in The Evolution of Mind: Fundamental Questions and Controversies, S.W. Gangestad and J. A. Simpson, eds. Guilford Publications, Inc., New York, NY.
  • 2007. R. Thornhill. Comprehensive knowledge of human evolutionary history requires both adaptationism and phylogenetics. Pp. 31-37 in The Evolution of Mind: Fundamental Questions and Controversies, S.W. Gangestad and J.A. Simpson, eds. Guilford Publications, Inc., New York, NY.
  • 2007. S.W. Gangestad and R. Thornhill. The evolution of social inference processes: The importance of signaling theory. Pp. 33-48 in Evolution and the Social Mind, J.P. Forgas, M.G. Haselton and W. von Hippell, eds. Psychology Press, New York, NY

  • 2006
  • 2006. C.E. Garver-Apgar, S.W. Gangestad, R. Thornhill, R. Miller and J. Olp. Major histocompatibility complex genes, sexual responsibility, and unfaithfulness in romantic couples. Psychological Science 17:830-835.
  • 2006. R. Thornhill. Foreword: Human sperm competition and woman’s dual sexuality. Pp. v-xvii in Sperm Competition in Humans: Classic and Contemporary Readings, T. K. Shackelford and N. Pound, eds. Springer-Verlag Press, New York, NY.
  • 2006. R. Thornhill and S.W. Gangestad. 2006. Facial sexual dimorphism, developmental stability and susceptibility to disease in men and women. Evolution and Human Behavior 27:131-144.
  • 2006 M. McIntyre, S.W. Gangestad, P.B. Gray, J.F. Chapman, T.C. Burnham, M.T. O’Rourke and R. Thornhill. Romantic involvement often reduces men’s testosterone levels–but not always: The moderating role of extrapair sexual interest. Journal of Personality and Social Psychology 91:642-651.

  • 2005
  • 2005. S.W. Gangestad, R. Thornhill and C.E. Garver-Apgar. Adaptations to ovulation: Implications for sexual and social behavior. Current Directions in Psychological Science 14:312-316.
  • 2005. S.W. Gangestad, R. Thornhill and C.E. Garver-Apgar. Women’s sexual interests across the ovulatory cycle depend on primary partner developmental instability. Proceedings of the Royal Society of London B 272:2023-2027.
  • 2005. S.W. Gangestad, R. Thornhill and C.E. Garver-Apgar. Adaptations to ovulation. Pp. 344-371 in The Handbook of Evolutionary Psychology, D.M. Buss, ed. John Wiley and Sons, Hoboken, NJ.
  • 2005. A.P. Møller, R. Thornhill and S.W. Gangestad. Direct and indirect tests for publication bias: Asymmetry and sexual selection. Animal Behaviour 70:497-506.

  • 2004
  • 2004. R. Thornhill and S.W. Gangestad. The evolution of human attractiveness and attraction. In Evolution: From Molecules to Ecosystems, A. Moya and E. Font, eds. Oxford University Press.
  • 2004. S.W. Gangestad and R. Thornhill. Female multiple mating and genetic benefits in humans: Investigations of design. In Sexual Selection in Primates, P. Kappler and C. van Schaik, eds. Cambridge University Press, Cambridge, UK.
  • 2004. R. Thornhill and C.T. Palmer. Evolutionary life history perspective on rape. Pp. 249-274 in Evolutionary Psychology, Public Policy and Personal Decisions, C. Crawford and C. Salmon, eds. Lawrence Erlbaum, Mahwah, NJ.

  • 2003
  • 2003. S.W. Gangestad and R. Thornhill. Facial masculinity and fluctuating asymmetry. Evolution and Human Behavior 24:231-241.
  • 2003. J. Radwan, P.J. Watson, J. Farslow, and R. Thornhill. Procrustean analysis of fluctuating asymmetry in the bulb mite, Rhizoglyphus robini Claparede (Astigmata: Acaridae). Biological Journal of the Linnean Society 80:499-505.
  • 2003. C.T. Palmer and R. Thornhill. Straw men and fairy tales: Evaluating reactions to A Natural History of Rape. Journal of Sex Research 40:249-255.
  • 2003. R. Thornhill, S.W. Gangestad, R. Miller, G. Scheyd, J. Knight and M. Franklin. MHC, symmetry, and body scent attractiveness in men and women. Behavioral Ecology 14:668-678.
  • 2003. R. Thornhill and C.T. Palmer. Rape and evolution: A reply to our critics. Psychology, Evolution and Gender 4:283-296.
  • 2003. K. Grammer, B. Fink, A.P. Møller and R. Thornhill. Darwinian aesthetics: Sexual selection and the biology of beauty. Biological Reviews 78:385-407.
  • 2003. R. Thornhill and S.W. Gangestad. Evolutionary theory leads to evidence for a male sex pheromone that signals symmetry. Psychological Inquiry 14:316-323.
  • 2003. C.T. Palmer and R. Thornhill. A posse of good citizens brings outlaw evolutionists to justice. A response to Evolution, Gender and Rape (2003, ed. by C.B. Travis, MIT Press, Cambridge, MA). Evolutionary Psychology 1:12-27.
  • 2003. R. Thornhill and S.W. Gangestad. Do women have evolved adaptation for extra-pair copulation? Pp. 341-368 in Evolutionary Aesthetics, K. Grammer and E. Voland, eds. Springer-Verlag, Berlin, Germany.
  • 2003. R. Thornhill. Darwinian aesthetics informs traditional aesthetics. Pp. 9-38 in Evolutionary Aesthetics, K. Grammer and E. Voland, eds. Springer-Verlag, Berlin, Germany.
  • 2003. C.T. Palmer and R. Thornhill. Rape and evolution: A reply to our critics. Psychology, Evolution and Gender 4:283-296.
  • 2003. S.W. Gangestad and R. Thornhill. Fluctuating asymmetry, developmental stability, and fitness: Toward a model-based interpretation. Pp. 62-80 in Developmental Instability: Causes and Consequences, M. Polak, ed. Oxford University Press, New York.
  • 2003. M. Polak, A.P. Møller, S.W. Gangestad, D.E. Kroeger, J.T. Manning and R. Thornhill. Does an individual asymmetry parameter exist? A meta-analysis. Pp. 81-98 in Developmental Instability: Causes and Consequences, M. Polak, ed. Oxford University Press, New York.

  • 2002
  • 2002. Gangestad, S.W., R. Thornhill and C. Garver. 2002. Changes in women’s sexual interest and their partners’ mate retention tactics across the menstrual cycle. Proceedings of the Royal Society of London B 269:975-982.
  • 2002. Kowner, R. and R. Thornhill. 2002. The imperfect organism: On the concept of asymmetry and its significance in humans, non-human animals and plants. Symmetry, Culture and Science 10:227-243.
  • 2002. K. Grammer, B. Fink, R. Thornhill, A. Juette, and G. Runzal. Female faces and bodies: N-dimensional feature space and attractiveness. In Facial Attractiveness: Evolutionary, Cognitive and Social Perspectives, G. Rhodes and L.A. Zebrowitz, eds. Greenwood Publishers Group, Westport, CT.

  • 2001
  • 2001. B. Fink, K. Grammer and R. Thornhill. Human (Homo sapiens) facial attractiveness in relation to skin texture and color. Journal of Comparative Psychology 115:92-99.
  • 2001. S. Gangestad, K. Bennett and R. Thornhill. A latent variable model of developmental instability in relation to men’s sexual behavior. Proceedings of the Royal Society of London B 268:1677-1684.
  • 2001. R. Thornhill and C.T. Palmer. New preface to paperback edition of A Natural History of Rape: Biological Bases of Sexual Coercion. Http://mitpress.mit.edu/thornhill-preface.html.
  • 2001. C.T. Palmer and R. Thornhill. Serial rape: An evolutionary perspective. Pp. 51-65 in Serial Offenders: Current Thought, Recent Findings, Unusual Syndromes, L.B. Schlesinger, ed. CRC Press.
  • 2001. C.T. Palmer, R. Thornhill and D.N. Dibari. Biology, sex and the debate over chemical castration. In Sexual Violence: Policies, Practice and Challenges. J.F. Hodgson and D.S. Kelly, eds. Praeger.

  • 2000
  • 2000. J.T. Manning, L. Barley, J. Walton, D.I. Lewis–Jones, R.L. Trivers, D. Singh, R. Thornhill, P. Rohde, T. Bereckei, P. Henzi, M. Soler and A. Szwed. The 2nd:4th digit ratio, sexual dimorphism, population differences and reproductive success: Evidence for sexually antagonistic genes. Evolution and Human Behavior 21:163-190.
  • 2000. R. Thornhill and C.T. Palmer. What A Natural History of Rape really says. The Albuquerque Tribune, March 2, pp. C1-2.
  • 2000. R. Thornhill and C.T. Palmer. What A Natural History of Rape really says. The Independent, February 21, p. 5.
  • 2000. R. Thornhill and C.T. Palmer. Just why do men rape, authors’ reply. The Sciences, pp. 6, 46-47.
  • 2000. R. Thornhill and C.T. Palmer. A Natural History of Rape: Biological Bases of Sexual Coercion. MIT Press.
  • 2000. R. Thornhill and C.T. Palmer. Why men rape. The Sciences 40:30-36.

  • 1999
  • 1999. R. Thornhill and S.W. Gangestad. Facial attractiveness. Trends in Cognitive Sciences 3:452-460.
  • 1999. R. Thornhill. The biology of human rape. Jurimetrics 39:137-147.
  • 1999. R. Thornhill. Sexual attraction, the evolutionary psychology of. Pp. 751-753 in The MIT Encyclopedia of the Cognitive Sciences, R.A. Wilson and F.C. Keil, eds. Boston, MA: MIT Press.
  • 1999. R. Thornhill and K. Grammer. The body and face of woman: One ornament that signals quality? Evolution and Human Behavior 20:105-120.
  • 1999. R. Thornhill and S.W. Gangestad. The scent of symmetry: A human sex pheromone that signals fitness? Evolution and Human Behavior 20:175-201.
  • 1999. R. Thornhill, A.P. Møller and S.W. Gangestad. The biological significance of fluctuating asymmetry and sexual selection: A reply to Palmer. American Naturalist 154:234-241.
  • 1999. A.P. Møller, S.W. Gangestad and R. Thornhill. Nonlinearity and the importance of fluctuating asymmetry as a predictor of fitness. Oikos 86:366-368.
  • 1999. J.E. Scheib, S.W. Gangestad and R. Thornhill. Facial attractiveness, symmetry and cues of good genes. Proceedings of the Royal Society of London, B, 266:1913-1918.
  • 1999. J. Manning, R. Trivers, D. Singh and R. Thornhill. The mystery of female beauty. Nature 399:214-215.
  • 1999. S.W. Gangestad and R. Thornhill. Individual differences in developmental precision and fluctuating asymmetry: A model and its implications. Journal of Evolutionary Biology 12:402-416.
  • 1999. R. Trivers, J. Manning, R. Thornhill, D. Singh and M. McGuire. The Jamaican asymmetry project: Long-term study of fluctuating asymmetry in rural Jamaican children. Human Biology 71:417-430.

  • 1998
  • 1998. A. Møller and R. Thornhill. Male parental care, differential parental investment by females and sexual selection. Animal Behaviour 55:1507-1515.
  • 1998. R. Thornhill and A.P. Møller. The relative importance of size and asymmetry in sexual selection. Behavioral Ecology 9:546-551.
  • 1998. G. Arnqvist and R. Thornhill. Evolution of animal genitalia: Patterns of phenotypic and genotypic variation and condition-dependence of genital and non-genital morphology in a water strider (Heteroptera: Gerridae). Genetical Research 71:193-212.
  • 1998. R. Thornhill and F. Bryant Furlow. Stress and human behavior: Attractiveness, women’s sexual development, post-partum depression, and baby’s cry. Pp. 319-369 in Advances in the Study of Behavior, P.J.B. Slater, M. Milinski and A. Møller, eds. Academic Press, New York, NY.
  • 1998. S.W. Gangestad and R. Thornhill. Menstrual cycle variation in women’s preferences for the scent of symmetrical men. Proceedings of the Royal Society of London, B, 265:927-933.
  • 1998. A. Møller and R. Thornhill. Developmental stability and sexual selection: A meta-analysis. American Naturalist 151:174-192.
  • 1998. S.W. Gangestad and R. Thornhill. The analysis of fluctuating asymmetry redux: The robustness of parametric statistics. Animal Behaviour 55:497-501.
  • 1998. R. Thornhill. Darwinian aesthetics. Pp. 543-572 in Handbook of Evolutionary Psychology: Ideas, Issues and Applications, C. Crawford and D. Krebs, eds. Lawrence Erlbaum Assoc., Mahwah, NJ.

  • 1997
  • 1997. R. Thornhill and A.P. Møller. Developmental stability, disease and medicine. Biological Reviews 72:497-548.
  • 1997. R. Thornhill. The concept of an evolved adaptation. Pp. 4-13 in Characterizing Human Psychological Adaptations, G. Bock and G. Cardew, eds. CIBA Foundation, London, UK.
  • 1997. G. Arnqvist, R. Thornhill and L. Rowe. Evolution of animal genitalia: Genital and non-genital morphology correlates of fitness components in a water strider. Journal of Evolutionary Biology 10:613-640.
  • 1997. A. Møller and R. Thornhill. Developmental instability is heritable: Reply. Journal of Evolutionary Biology 10:69-70.
  • 1997. F.B. Furlow, T. Armijo–Prewitt, S.W. Gangestad and R. Thornhill. Psychometric intellegience and fluctuating asymmetry. Proceedings of the Royal Society, London, B, 264.
  • 1997. S.W. Gangestad and R. Thornhill. The evolutionary psychology of extra-pair sex: The role of fluctuating asymmetry. Evolution and Human Behavior 18:69-88.
  • 1997. S.W. Gangestad and R. Thornhill. Human sexual selection and developmental stability. Pp. 169-195 in Evolutionary Social Psychology, J.A. Simpson and D.T. Kenrick, eds. Lawrence Erlbaum Assoc., Mahwah, NJ.
  • 1997. A. Møller and R. Thornhill. A Meta-analysis of the heritability of fluctuating asymmetry. Journal of Evolutionary Biology 10:1-16.
  • 1997. R. Thornhill. Rape-victim psychological pain revisited. Pp. 239-249 in Evolution and Human Behavior: A Critical Reader, Laura Betzig, ed. Oxford University Press.
  • 1997. J.T. Manning, R.L. Trivers, R. Thornhill, D. Singh, J. Denman, M.H. Eklo, and R.H. Anderton. Ear asymmetry and left-side cradling. Evolution and Human Behavior 18:327-340.

  • 1996
  • 1996. R. Thornhill and S.W. Gangestad. Human female copulatory orgasm: A human adaptation or phylogenetic legacy? Animal Behaviour 52:853-855.
  • 1996. R. Thornhill and S.W. Gangestad. The evolution of human sexuality. Trends in Ecology and Evolution 11:98-102.
  • 1996. B. Furlow and R. Thornhill, The orgasm wars. Psychology Today 29:42-46.

  • 1995
  • 1995. A.P. Møller, M. Soler and R. Thornhill. Breast asymmetry, sexual selection and human reproductive success. Ethology and Sociobiology 16:207-219.
  • 1995. R. Thornhill, S.W. Gangestad and R. Comer. Human female orgasm and mate fluctuating asymmetry. Animal Behaviour 50:1601-1615.

  • 1994
  • 1994. R. Thornhill and S.W. Gangestad. Fluctuating asymmetry correlates with lifetime sex partner numbers and age at first sex in Homo sapiens. Psychological Science 5:297-302.
  • 1994. S. Gangestad, R. Thornhill and R. Yeo. Facial physical attractiveness, developmental stability and fluctuating asymmetry. Ethology and Sociobiology 15:73-85.
  • 1994. R. Thornhill. Is there psychological adaptation to rape? Analyse and Kritik 16:68-85.
  • 1994. P. Watson and R. Thornhill. Fluctuating asymmetry and sexual selection. Trends in Ecology and Evolution 9:21-24.
  • 1994. K. Grammer and R. Thornhill. Human (Homo sapiens) facial attractiveness and sexual selection: The role of symmetry and averageness. Journal of Comparative Psychology 108:233-242.

  • 1993
  • 1993. R. Thornhill and S. Gangestad. Human facial beauty: Averageness, symmetry and parasite resistance. Human Nature 4:237-269.
  • 1993. K. Johnson, R. Thornhill, J.D. Ligon and M. Zuk. The directions of mothers’ and daughters’ preferences and the heritability of male ornaments in red jungle fowl (Gallus gallus). Behavioral Ecology 4:254-259.
  • 1993. R. Thornhill. The allure of symmetry. Natural History 102:30-37.

  • 1992
  • 1992. M. Zuk, J.D. Ligon and R. Thornhill. Effects of experimental manipulation on male secondary sex characters on female mate preference in red jungle fowl. Animal Behaviour 44:999-1006.
  • 1992. R. Thornhill. Fluctuating asymmetry and the mating system of the Japanese scorpionfly, Panorpa japonica. Animal Behaviour 44:867-879.
  • 1992. R. Thornhill and N.W. Thornhill. The study of men’s coercive sexuality: What course should it take? Behavioral and Brain Sciences (authors’ response to 28 commentators) 15:404-421.
  • 1992. R. Thornhill and N.W. Thornhill. The evolutionary psychology of men’s sexual coercion. Behavioral and Brain Sciences (target article) 15:363-375.
  • 1992. R. Thornhill. Fluctuating asymmetry and interspecific competition in two species of Japanese scorpionflies. Behavioral Ecology and Sociobiology 30:357-363.
  • 1992. R. Thornhill. Females prefer the pheromone of males with low fluctuating asymmetry in the Japanese scorpionfly (Panorpa japonica). Behavioral Ecology 3:277-283.
  • 1992. R. Thornhill and K.P. Sauer. Paternal genetic effects on the fighting ability of sons and daughters and mating success of sons in a scorpionlfy (Panorpa vulgaris). Animal Behavior 43:255-264.

  • 1991
  • 1991. S. Austad and R. Thornhill. This bug’s for you. Natural History (No. 12.):44-49.
  • 1991. R. Thornhill. Teleonomy and the study of sexual selection. Acta XX Congressus Internationalis Ornitholgici, Vol. III:1361-1366.
  • 1991. N.W. Thornhill and R. Thornhill. Evolutionary analysis of psychological pain following rape IV: The effect of the nature of the sex act. J. Comp. Psychology 105:243-252.
  • 1991. R. Thornhill and K.P. Sauer. The notal organ of the scorpionfly (Panorpa vulgaris): An adaptation to coerce mating duration. Behavioral Ecology 2:156-164.

  • 1990
  • 1990. M. Zuk, R. Thornhill, J.D. Ligon and K. Johnson. Parasites and mate choice in red jungle fowl. American Zoologist 30:235-244.
  • 1990. R. Thornhill and N.W. Thornhill. Coercive sexuality of men: Is there psychological adaptation to rape? In Sexual Coercion: Its Nature, Causes and Prevention, E.Grauerholz and M. Koralewski, eds. Lexington Books.
  • 1990. M. Zuk, K. Johnson, R. Thornhill and J.D. Ligon. Female red jungle fowl show a threshold mate preference. Evolution 44:477-485.
  • 1990. J.D. Ligon, R. Thornhill, M. Zuk, and K. Johnson. Male–male competition and male ornamentation in red jungle fowl. Animal Behavior 40:367-373.
  • 1990. R. Thornhill. The study of adaptation. Pp. 31-62 in Interpretation and Explanation in the Study of Behavior, Vol. II, M. Bekoff and D. Jamieson, eds. Westview Press.
  • 1990. N.W. Thornhill and R. Thornhill. Evolutionary analysis of psychological pain of rape victims I: The effects of victim’s age and marital status. Ethology and Sociobiology 11:155-176.
  • 1990. N.W. Thornhill and R. Thornhill. Evolutionary analysis of psychological pain following rape II: The effects of stranger, friend and family member offenders. Ethology and Sociobiology 11:177-193.
  • 1990. N.W. Thornhill and R. Thornhill. Evolutionary analysis of psychological pain following rape III: The effects of force and violence. Aggressive Behavior 16:297-320.
  • 1990. M. Zuk, K. Johnson, R. Thornhill and J.D. Ligon. Parasites and sexual selection in red jungle fowl. Behaviour 114:232-248.
  • 1990. M. Zuk, R. Thornhill, J.D. Ligon, K. Johnson, S. Ligon, N.W. Thornhill, S. Austad and C. Costin. Female choice and male ornamentation in red jungle fowl. American Naturalist 136:459-473.

  • 1989
  • 1989. R. Thornhill and N.W. Thornhill. The evolution of psychological pain. Pp. 73-103 in Sociobiology and the Social Sciences, R. Bell, ed. Texas Tech Univ. Press.
  • 1989. R. Thornhill. Nest defense by red jungle fowl (Gallus gallus spadiceus) hens: The roles of renesting potential, parental experience and brood reproductive value. Ethology 83:31-42.
  • 1989. R. Montgomerie and R. Thornhill. Fertility advertisement in birds: A means of inciting male–male competition. Ethology 81:209-220.

  • 1988
  • 1988. R. Thornhill. The hen’s cackle incites male competition. Veh. Dtsch. Zool. Ges. 81:145-154.

  • 1987
  • 1987. N.W. Thornhill and R. Thornhill. Female primate sexual behavior and conception: Are there really sperm to spare? Current Anthropology 29:93-94.
  • 1987. L. Heisler, M.B. Andersson, S.J. Arnold, C.R. Boake, G. Borgia, G. Hausfater, M. Kirkpatrick, R. Lande, J. Maynard Smith, P. O’Donald, R. Thornhill and F.J. Weissing. The evolution of mating preferences and attractive traits. In Sexual Selection: Testing the Alternatives, J. Bradbury and M. Andersson, eds. Springer–Verlag, New York, NY.
  • 1987. N.W. Thornhill and R. Thornhill. Evolutionary theory and rules of mating and marriage pertaining to relatives. Pp. 373-400 in Psychology and Sociobiology: Ideas, Issues, and Applications, C. Crawford, M. Smith, and D. Krebs, eds. Erlbaum Assoc. Publ., London.
  • 1987. R. Thornhill and N.W. Thornhill. Human rape: the strengths of the evolutionary perspective. Pp. 269-291 in Psychology and Sociobiology: Ideas, Issues and Applications, C. Crawford, M. Smith and D. Krebs, eds. Erlbaum Assoc. Publ., London.
  • 1987. R. Thornhill. The relative importance of intra- and interspecific competition in scorpionfly mating systems. Am. Nat. 130:711-729.

  • 1986
  • 1986. R. Thornhill, N.W. Thornhill and G. Dizinno. The biology of rape. In Rape, S. Tomaseli and R. Porter, eds. Oxford, England: Basil Blackwell and Welcome Institute for the History of Medicine.
  • 1986. S. Austad and R. Thornhill. Female reproductive variation in a nuptial-feeding spider, Pisaura mirabilis. Bull. Br. Arachnol. Soc. 7:48-52.
  • 1986. R. Thornhill and D. Gwynne. The evolution of sexual differences in insects. Amer. Scientist 74:382-389.
  • 1986. R. Thornhill. Relative parental contribution of the sexes to their offspring and the operation of sexual selection. Pp. 113-136 in Evolution of Animal Behavior, M. Nitecki and J. Kitchell, eds. Oxford: Oxford Univ. Press.

  • 1985
  • 1985. N.W. Thornhill and R. Thornhill. Matriliny and sexual selection and conflict. Behav. and Brain Sciences 8:679-680.

  • 1984
  • 1984. R. Thornhill. Alternative hypotheses for traits believed to have evolved by sperm competition. Pp. 151-178 in Sperm Competition and the Evolution of Animal Mating Systems, R.L. Smith, ed. Academic Press, New York, NY.
  • 1984. R. Thornhill. Alternative female choice tactics in the scorpionfly Hylobittacus apicalis (Mecoptera) and its implications. Amer. Zoologist. 24:367-383.
  • 1984. R. Thornhill. Fighting and assessment in Harpobittacus scorpionflies (Insecta: Mecoptera). Evolution 38:204-214.
  • 1984. R. Thornhill. Scientific methodology in entomology. Florida Entomol. 67:74-96.

  • 1983
  • 1983. R. Thornhill. Cryptic female choice in the scorpionfly Harpobittacus nigriceps and its implications. American Naturalist 122:765-788.
  • 1983. R. Thornhill and N.W. Thornhill. Human rape: An evolutionary analysis. Ethology and Sociobiology 4:137-173.
  • 1983. R. Thornhill, G. Dodson and L. Marshall. Sexual selection and insect mating behavior. American Biology Teacher 45:310-319.
  • 1983. G. Byers and R. Thornhill. Biology of the Mecoptera. Ann. Review Entomology 28:203-228.

  • 1982
  • 1982. W. Shields, L. Shields, R. Thornhill and N.W. Thornhill. What causes rape? A dissenting view. Science 82 (Dec.):16.

  • 1981
  • 1981. R. Thornhill. Panorpa (Mecoptera: Panorpidae) scorpionflies: Systems for understanding resource-defense polygyny and alternative male reproductive efforts. Ann. Rev. Ecol. and Syst. 12:355-386.

  • 1980
  • 1980. R. Thornhill. Sexual selection in the black-tipped hangingfly. Scientific American 242:162-172.
  • 1980. R. Thornhill. Competitive, charming males and choosy females: Was Darwin correct? Florida Entomologist 63:5-30.
  • 1980. R. Thornhill. Mate choice in Hylobittacus apicalis (Insecta: Mecoptera) and its relation to some models of female choice. Evolution 34:519-538.
  • 1980. R. Thornhill. Sexual selection within mating swarms of the lovebug, Plecia nearctica (Diptera: Bibionidae). Animal Behavior 28:405-412.
  • 1980. R. Thornhill. Rape in Panorpa scorpionflies and a general rape hypothesis. Animal Behavior 28:52-59.
  • 1980. R. Thornhill. Competition and coexistence in Panorpa scorpionflies (Mecoptera: Panorpidae). Ecological Monographs 50:179-197.

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