1966. B.S., Biology, Math, Secondary Education, University of Wisconsin-River Falls, River Falls, WI.
1968. M.S., Zoology, Colorado State University, Ft. Collins, CO. Thesis title: Eimeria of the cottontail rabbit, Sylvilagus audubonii, in northeastern Colorado.
Major Advisor: W.C Martquardt.
1970. Ph.D., Zoology, Colorado State University, Ft. Collins, CO. Dissertation title: Interspecific interaction between two species of Coccidia (Sporozoa: Eimeriidae) in the rat.
Major Advisor: W.C. Martquardt.
Other colleges attended:
University of Costa Rica, San Jose, Costa Rica. Summer, 1967.
University of the Pacific, Stockton, CA. Summer, 1968.
Vanderbilt University, Nashville, TN. Summer, 1972.
University of Oklahoma, Lake Texoma, Summer, 1998.
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