Neurophysiology/Neuroanatomy (Biomed 533)

Fall 2009

Lectures (rm 243 BMSB): Tues & Thurs 8:30 – 10:00

Labs (rm 382): Thurs 2:00 – 4:00

http://www.unm.edu/~neurohsc/course533.htm

 

Note:  All students must complete the ARF Animal Use Certification before 12 November.

Paper Assignments

 

Date

Topic

Reading

Assigned Paper

Faculty

Tu 25 Aug

Electrophysiology review

5,6 S

 

 

Partridge

Th 27 Aug

Surface anatomy of the brain / meninges / blood supply

2 S

3,4 MB

 

Cunningham

Th 27 Aug

General structure of the Brain / Meninges

3,4 MB

 

Cunningham

Tu 1 Sept

Pathways of special senses

 

Palmer, 2009 (25 Aug)

Saland

Th 3 Sept

Sensory processing

23 S

 

Partridge

Th 3 Sept

Blood supply

5 MB

 

Cunningham

Tu 8 Sept

Anatomy & physiology of retina (trichromacy evolution)

27 S

Glazewski, 2007 (3 Sept)

Partridge

Th 10 Sept

Central visual physiology

27 S

Roska, 2006 (8 Sept)

Partridge

Th 10 Sept

Topology of the cerebral hemispheres

6 MB

 

Cunningham

Tu 15 Sept

Physiology of taste

24 S

Ohki, 2005 (10 Sept)

Partridge

Th 17 Sept

Olfactory physiology

24 S

Maruyama, 2006 (15 Sept)

Partridge

Th 17 Sept

Quiz 1  3D anatomy of internal structures movie

 

 

Cunningham

Tu 22 Sept

Auditory physiology

26 S

Oka, 2006 (17 Sept)

Partridge

Th 24 Sept

BSGP Graduate Retreat

 

Tu 29 Sept

Somatosensory physiology

25 S

 

Partridge

Th 1 Oct

Sensory & motor pathways

23, 28 S

Dallos, 2008 (22 Sept)

Saland

Th 1 Oct

White matter of cerebral hemispheres / basal ganglia)

7 MB

 

Cunningham

Tu 6 Oct

Proprioception

25 S

Ferezou, 2006 (29 Sept)

Partridge

Th 8 Oct

Spinal Reflexes

29 S

Albert, 2006 (6 Oct)

Partridge

Th 8 Oct

Lateral ventricles and rhinencephalon

8 MB

 

Cunningham

Tu 13 Oct

Postural Mechanisms

30 S

Xia, 2005 (8 Oct)

Partridge

Th 15 Oct

Fall Break

 

 

 

Tu 20 Oct

SfN meeting

 

 

 

Th 22 Oct

Limbic System Anatomy

 

Lakie, 2005 (13 Oct)

Wallace

Th 22 Oct

Limbic System

9MB

 

Cunningham

Tu 27 Oct

Locomotion (evolution of endurance running)

29 S

 

Yakovenko, 2005 (27 Oct)

Partridge

Th 29 Oct

Cerebellar Anatomy

32 S

 

Cunningham

Th 29 Oct

Brainstem, cranial nerves, cerebellum

10, 11 MB

 

Cunningham

Tu 3 Nov

Oculomotor function

33 S

Hu, 2007 (3 Nov)

Partridge

Tu 5 Nov

Voluntary control of movement

30 S

Velliste 2008 (5 Nov)

Partridge

Th 5 Nov

Quiz 2

 

 

Cunningham

Tu 10 Nov

Autonomic Physiology

35 S

 

Shuttleworth

Tu 12 Nov

Sleep and Circadian Rhythms

41, 42 S

He, 2009 (12 Nov)

Partridge

Th 12 Nov

Rodent brain structures

 

 

Cunningham

Tu 17 Nov

Motivation and Reward

43 S

 

Valenzuela

Th 19 Nov

Rodent brain structures (atlas)

 

 

Cunningham

Th 19 Nov

Project Handbook

Lagrace, 2007

 

Cunningham

Tu 24 Nov

Project

 

 

Cunningham

Th 26 Nov

Thanksgiving

 

 

 

Tu 1 Dec

Learning and Memory I

49 S

Ranganath, 2004 (1 Dec)

Wilson

Th 3 Dec

Learning and Memory II

50 S

Roberts, 2009, Hairston, 2005 (3 Dec)

Perrone

Th 3 Dec

Project

Cunningham

Tu 8 Dec

Quiz 3

 

 

Cunningham/Partridge

Th 10 Dec

Consciousness

Leutgeb, 2005

 

Partridge

Th 10 Dec

Project Discussions

 

 

Cunningham/Partridge

 

Grading

Paper presentations 30%

Project 30%

            20% Written paper

            10% Oral discussion

Anatomy Quizzes 30%

Class participation 10%

Reading Assignments

Reading assignments are given as chapters in: 

S Squire, L R. et al. Fundamental Neuroscience 3rd ed. Elsevier, 2008.

M - Montemurro & Bruni The Human Brain in Dissection 2nd ed. Oxford University Press

Additional Assigned readings

Leutgeb, S., Leutgeb, J.K., Moser, M., Moser, E.I. (2005) Place cells, spatial maps and the population code for memory.  Current Opinion in Neurobiol. 15:109

This paper discusses the evidence for an attractor-based population code for memory based in the recurrent networks of the hippocampus and entorhinal cortex. 

Lagrace et al., 2007, Dynamic contribution of nestin-expressing stem cells to adult neurogenesis. J. Neuroscience 14:12623-12629.

 

 

Project – Histological methods for identifying multipotent stem cells in the CNS.  Students will complete a genetic fate mapping study of neural stem cells within the adult mouse CNS using an inducible Nestin-Cre-loxP mouse (see Lagace et al., 2007, J. Neuroscience 14; 12623 for description of this mouse line)  Students will participate in all aspects of tissue prep, histological sectioning of tissue and immunostaining using various antibodies directed against stem cells and/or newly born neurons in hippocampus subgranular zone and subventricular zones lining the lateral ventricles.  Students will photodocument their results and include them in the written paper (described below).  The results will be discussed in the larger context of adult stem cells in the CNS during the Project Discussions (see below).

1. Paper

Each student will write a 10 page focused paper that discusses some aspect of  Stem cell function in the adult nervous system. Students should choose the precise topic after discussion with the course faculty by Tuesday, 3 September.  Papers are due on Thursday, 24 November. Some examples include: Electrophysiological properties of adult born granule cell neurons, Mechanisms of functional incorporation of adult born neurons into existing circuitries, Role of adult neurogenesis in memory, Injury induced neurogenesis in brain and spinal cord, Therapeutic potential of neural stem cells in degenerative disease….etc.. The paper should include the following:

1. Introduction (1-2 pages) that briefly discusses the topic background 2. Methods (1-2 pages) that were used to identify neural stem cells or new born neurons in your project..

3. Discussion (6-8 pages) of the most important findings in the chosen topic area 4. Future Directions: What needs to be done to advance the field to the next level.

5. References.

The paper should be clearly written using the format of any of the papers on the assigned reading list.  The inclusion of appropriate (referenced) figures is encouraged and these will not be included in the page limits.

2. Project Discussions

All students should be prepared to discuss their projects during a 2 hr session on Dec 10.  This discussion may include:  strengths and pitfalls of methods used, additional potential applications of the technique, and specific details of the functional and behavioral correlates examined in the student’s paper.  Students will be graded in part on their scholarly participation in this discussion.

 

Assigned Paper Discussions

The papers listed with certain topics will be the basis of an in-depth discussion during the first half hour of class following the in-class discussion of that topic.  (Dates following papers refer to the date when that topic was covered in lecture.)  All students should read and be prepared to discuss the assigned paper.  The student presenting may need to read additional papers in order to put the assigned paper into context and lead a discussion of the paper.  Discussions should include the following:

·  about 10 minute introduction to the paper including sufficient methods to understand the paper

·  about 20 minutes of discussion concentrating on the figures and conclusions of the paper

 

References for assigned paper presentations

Albert, F., Bergenheim, M. Ribot-Ciscar, E.R., Roll, J.-P. (2006) The Ia afferent feedback of a given movement evokes the illusion of the same movement when returned to the subject via muscle tendon vibration.  Exp. Brain Res. 172:163-174.

In addition to providing the afferent limb of the stretch reflexes, muscle spindles are the sensors for conscious proprioception.  In this unusual study, Ia activity from six main muscle groups on the ankle were recorded during a “writing like” movement.  At a later time, this pattern was “played back” into the spindles and the subjects reported illusionary perception of the original movement.

Dallos, P., Wu, X., Cheatham, M.A., Gao, J., Zheng, J. Anderson, C.T., Jia, S., Want, X., Chang, W.H., Senqupta, S., He, D.Z., Zuo, J. (2008) Prestin-based outer hair cell motility is necessary for mammalian cochlear amplification.  Neuron 58:333 – 339.

This study uses a knockout of the outer hair cell protein prestin.  The results suggest that the presence of prestin is a requirement for coupling mechanical vibrations between the basilar membrane and stereocillia.  Furthermore, the results that sharp tuning of frequency on the basilar membrane requires amplification by the outer hair cells.

 

Ferezou, I., Bolea, S., Petersen, C.C.H. (2006) Visualizing the cortical representation of whisker touch:  Voltage-sensitive dye imaging in freely moving mice.  Neuron 50:617-629.

In another technical tour de force, this paper uses optical recording of voltage sensitive dyes to permit measurements of the activity of somatosensory cortex activity in freely moving mice.  They show that the neural response to whisking is dependent on the behavior of the animal.

Glazewski, S., Benedetti, B.L., Barth, A.L., (2007) Ipsilateral whiskers suppress experience-dependent plasticity in the barrel cortex. J. Neurosci. 27:3910-3920

This is a study of the effect of converging information from both sides of the body onto receptive fields in the somatosensory cortex.  Individual whiskers in the rat map onto discrete cortical barrel columns, which can be deprived of their input by simply shaving a whisker.  This paper demonstrates that the size of somatosensory receptive fields depends on both contralateral and ipsilateral sensory input.

Hairston, Little, Scanlon, Barakat, Palmer, Sapolsky, Heller (2005) Sleep restriction suppresses neurogenesis induced by hippocampus-dependent learning.  J. Neurophysiol

Fully rested animals use a spatial strategy even when this strategy interferes with performance in a non-spatial task. Conversely, sleep restricted animals used non-spatial information, and hence improve performance in a non-spatial task.

He, Y., Jones, C.R., Fujiki, N., Xu, Y., Guo, B., Holder, J.L., Rossner, M.J., Fu, Y-H. (2009) The transcriptional repressor DEC2 regulates sleep length in mammals. Science 325:866-870 (introductory material)
Humans need from less than 6 to more than 9 hours of sleep, but short sleepers and long sleepers appear to run in families. This study uses activity profiles and sleep recordings from humans, mice, and drosophila to identify a mutation in a transcriptional factor DEC2 that is associated with short sleep.

Hu, X., Jiang, H., Gu, C., Li, C. Sparks, D.L. (2007) Reliability of oculomotor command signals carried by individual neurons.  PNAS 104:8137-8142

Motor commands for saccadic eye movements are produced by bursts of activity in neurons of the pontine reticular formation.  This study shows that high temporal precision of the firing of these neurons underlies the accuracy and precision of the eye movements.

Lakie, M., Loram, I.D., (2006) Manually controlled human balancing using visual, vestibular and proprioceptive senses involves a common, low frequency neural process. J. Physiol. 577:403-416

Subjects used visual, vestibular, or proprioceptive sensation to balance their own body or a mechanically equivalent unstable inverted pendulum.  From these data the authors conclude that the control of standing  requires a common central motor planning process.

 

Maruyama, Y., Pereira, E., Margolskee, R.F., Chaudhari, N., Roper, S.D., (2006) Umami responses in mouse taste cells indicate more than one receptor.  J. Neurosci. 26:2227-2234

In addition to the traditional 4 tastes of sour, sweet, salty, and bitter, there is an additional sensory dimension – umami or the taste of L-glutamate.  These authors conclude that this sensation must result from novel combinations of receptors, some of which have not yet been discovered.

 

Ohki, K., Chung, S., Ch’ng, Y.H., Kara, P. Reid, R.C. (2005) Functional imaging with cellular resolution reveals precise microarchitecture in visual cortex. Nature 433:597-603

The arrangement of the visual cortex into vertical columns of neurons with similar orientation selectivity was one of the seminal findings in the classic work of Hubel and Weisel.  This study uses Ca2+ imaging and two-photon microscopy to determine the precision of single cells in this exquisite organization.

Oka, Y., Katada, S., Omura, M., Suwa, M., Yoshihara, Y., Touhara, K. (2006) Odorant receptor map in the mouse olfactory bulb:  In vivo sensitivity and specificity of receptor-defined glomeruli. Neuron 52: 857-869.

Odor discrimination results from mapping odorants in a complex sensory space of odorant receptors and glomerular activity.  In order to define the first step in this process this study used Ca2+ imaging, retrograde dye labeling, and single-cell RT-PCR to identify odorant receptors.  This is a first step in understanding olfactory specificity under physiological conditions.

Palmer, L.M., Stuart, G.J. (2009) Membrane potential changes in dendritic spines during action potentials and synaptic input. J. Neurosci. 29:6897-6903

Excitatory input to neurons occurs at dendritic spines, but the electrical properties of these spines is not well characterized.  The authors of this study used voltage-sensitive dyes to measure the electrical signals in individual spines during backpropagated action potentials and synaptic input.  They conclude that spines behave passively and do not significantly modify the synaptic inputs.

Ranganath, C., Cohen, M.X., Dam, C., D'Esposito, M. (2004) Inferior Temporal, Prefrontal, and HippocampalContributions to Visual Working Memory Maintenance and Associative Memory Retrieval. J. Neurosci. 24:3917- 3925

This paper examines the idea that higher order cognition depends on the ability to recall information and hold it in mind to guide future behavior and that these functions may be encoded in different brain regions. The study uses fMRI analysis to compare brain activity during performance of a visual associative and working memory tasks. fMRI provides an indirect assay of fairly localized brain activity, the student is encouraged not to discuss fMRI methodology in great detail, but focus on the broader physiological implications of the study.

Roberts, G.M.P, Nestor, L., Garavan, H. (2009) Learning and memory deficits in ecstacy users and their neural correlates during a face-learning task.  Brain Res. Epub.

This study uses fMRI to address the memory dysfunction of ecstasy users with a modified version of the face-name task.  The study found decreaed activity in the right medial frontal gyrus, left parahippocampal gyrus, left dorsal cingulated gyrus, and left caudate.

 

Roska, B., Molnar, A., Werblin, F.S., (2006) Parallel processing in retinal ganglion cells: How integration of space-time patterns of excitation and inhibition form the spiking output.  J. Neurophysiol. 95:3810-3822

Complex processing of visual information occurs at the level of the retina, which has receptors, 3 classes of interneurons, and output ganglion neurons.  This study correlates morphological studies of ganglion cells with space-time maps of their firing patterns.

Velliste, M., Perel, S., Spaling, M.C., Whitford, A.S., Schwartz, A.B. (2008) Cortical control of a prosthetic arm for self-feeding.  Nature 453:1098-1101

Monkeys rapidly learn a brain-machine-interface that uses electrodes implanted in the motor cortex to control a prosthetic device to feed themselves. 

Xia, R., Bush, B.M.H., Karst, G.M. (2005) Phase-dependent and task-dependent of stretch reflexes during rhythmic hand tasks in humans. J. Physiol. 564:941-951.

Reflexes were evoked by mechanical perturbations at various phases of tasks akin to handwriting.  Stretch reflexes were found to be modulated with respect to the phase of the rhythmic task.  It was concluded that central modulation of hand reflexes reflects optimal modulation strategies.

 

Yakovenko, S., McCrea, D.A., Stecina, K., Prochazha, A. (2005) Control of locomotor cycle durations.  J. Neurophysiol. 94:1057-1065.

The rate of locomotor cadence varies as a result of flexor rather than extensor phase durations.  This behavior could be modeled with a simple half-center model of the locomotor central pattern generator.